© Charlie Christensen


Departamental II – DII. 211
Area of Biodiversity and Conservation
Universidad Rey Juan Carlos
c/ Tulipán s/n.
E-28933 Móstoles (Madrid)

Phone: +34 91 488 8249
Fax:       +34 91 664 7490
E-mail: marcos.mendez at urjc.es

Short CV

1990 BsC Biology, Zoology – Oviedo University (Spain)
1996 PhD Thesis – Oviedo University (Spain)
1997-1998 Post-doc fellowship – Uppsala University (Sweden)
1999-2001 Post-doc fellowship – Uppsala University (Sweden)
2002-2003 Assistant lecturer – Stockholm University (Sweden)
2004-2009 Assistant lecturer – Rey Juan Carlos University (Spain)
2010- Associate professor (senior lecturer) – Rey Juan Carlos University (Spain)

Current position

Associate professor (senior lecturer).

I teach Evolutionary Biology in the degrees of Biology and Experimental Sciences.

I am the coordinator of the doctoral program in Conservation of Natural Resources.

Research interests

Plant evolutionary ecology:
– Resource economy and cost of reproduction.
– Gamete packaging and sexual expression.
– Pollination and evolution of floral morphology.

Useful indices in plant reproductive ecology.

Community ecology:
– Community structure in arthropods.


53. Moncalvillo, B.; Méndez, M.; Iriondo, J. M. (In press). Ecotypic differentiation reveals seed-color-related alkaloid contents in a crop wild relative. Plant Biology.

52. Torices, R.; Afonso, A.; Anderberg, A. A.; Gómez, J. M.; Méndez, M. (2019). Architectural traits constrain the evolution of unisexual flowers and sexual segregation within inflorescences: an interspecific approach. bioRxiv, 356147, ver. 3, peer-reviewed and recommended by PCI Evolutionary Biology.

51. Herrera, P.; Suárez, J. P.; Sánchez-Rodríguez, A.; Molina, M. C.; Prieto, M.; Méndez, M. (2019). Many, broadly shared mycobionts characterize mycorrhizal interactions of two coexisting epiphytic orchids in a high elevation tropical forest. Fungal Ecology 39: 26-36.

50. Méndez, M. (2018). Ten simple rules for developing good reading habits during graduate school and beyond. PLoS Computational Biology 14: e1006467. PDF     Access the recommendation on F1000Prime

49. Cayuela, L.; Granzow de la Cerda, I.; Méndez, M. (2018). The state of European research in tropical biology. Biotropica 50: 202-207. PDF

48. Jara-Guerrero, A. K.; Escribano-Ávila, G.; Espinosa, C.; de la Cruz, M.; Méndez, M. (2018). Deer as the last megafauna dispersing seeds in Neotropical dry forests: the role of fruit and seed traits. Biotropica 50: 169-177.

47. Herrera, P.; Kottke, I.; Molina, M. C.; Méndez, M.; Suárez, J. P. (2018). Generalism in the interaction of Tullasnellaceae mycobionts with orchids characterizes a biodiversity hotspot in the tropical Andes of Southern Ecuador. Mycoscience 59: 38-48.

46. Méndez, M.; de Jaime, C.; Alcántara, M. A. (2017). Habitat description and interannual variation in abundance and phenology of the endangered beetle Lucanus cervus L. (Coleoptera) using citizen science monitoring. Journal of Insect Conservation 21: 907-915.

45. Thomaes, A.; Verschelde, P.; Mader, D.; Sprecher-Uebersax, E.; Fremlin, M.; Onkelinx, T.; Méndez, M. (2017). Can we successfully monitor population density decline of elusive invertebrates? A statistical power analysis on Lucanus cervus. Nature Conservation 19: 1-18.

44. Merinero, S.; Méndez, M.; Aragón, G.; Martínez, I. (2017). Variation in the reproductive strategy of a lichenized fungus along a climatic gradient. Annals of Botany 120: 63-70.

43. Granado-Yela, C.; Balaguer, L.; Cayuela, L.; Méndez, M. (2017). Unusual positional effects on flower sex in an andromonoecious tree: resource competition, architectural constraints or inhibition by the apical flower? American Journal of  Botany 104: 608-615.

42. Campanaro, A.; Zapponi, L.; Hardersen, S.; Méndez, M.; Al Fulaij, N.; Audisio, P.; Bardiani, M.; Carpaneto, G. M.; Corezzola, S.; Della Rocca, F.; Harvey, D.; Hawes, C.; H.; Kadej, M.; Karg, J.; Minari, E.; Rink, M.; Smolis, A.; Sprecher, E.; Thomaes, A.; Toni, I.; Vrezec, A.; Zauli, A.; Zilioli, M.; Chiari, S. (2016). A European monitoring protocol for the stag beetle, a saproxylic flagship species. Insect Conservation and Diversity 9: 574-584. PDF

41. Gómez, J. M.; Verdú, M.; González-Mejías, A.; Méndez, M. (2016). The phylogenetic roots of human lethal violence. Nature 538: 233-237. PDF

40. Valencia, E.; Méndez, M.; Saavedra, N.; Maestre, F. T. (2016). Plant size and leaf area influence phenological and reproductive responses to warming in semiarid Mediterranean species. Perspectives in Plant Ecology, Evolution and Systematics 21: 31-40. PDF

39. Santamaría, S.; Galeano, J.; Pastor, J. M.; Méndez, M. (2016). Removing interactions, rather than species, casts doubt on the high robustness of pollination networks. Oikos 125: 526-534. PDF

38. Álvarez-Lao, D.; Méndez, M. (2016). Latitudinal gradients and indicator species in ungulate paleoassemblages during the MIS 3 in W Europe. Palaeogeography, Palaeoclimatology, Palaeoecology 449: 455-462. PDF

37. Méndez, M.; Escudero, A.; Iriondo, J. M.; Viejo, R. (2015). Demography gone wild in native species: four reasons to avoid the term “native invader”. Web Ecology 14: 85-87. PDF

36. Jara-Guerrero, A.; de la Cruz, M.; Espinosa, C. I.; Méndez, M.; Escudero, A. (2015). Does spatial heterogeneity blur the signature of dispersal syndromes on spatial patterns of woody species? A test in a tropical dry forest. Oikos 124: 1360-1366. PDF

35. Santamaría, S.; Galeano, J.; Pastor, J. M.; Méndez, M. (2014). Robustness of alpine networks: effects of network structure and consequences for endemic plants. Arctic, Antarctic and Alpine Research 46: 568-580. PDF

34. Torices, R.; Méndez, M. (2014). Resource allocation to inflorescence components is highly integrated despite differences between allocation currencies and sites. International Journal of Plant Sciences 175: 713-723. PDF

33. Granado-Yela, C.; Balaguer, L.; García-Verdugo, C.; Carrillo, K.; Méndez, M. (2013). Triving at the limit: Differential reproductive performance in range-edge populations of a Mediterranean sclerophyll (Olea europaea). Acta Oecologica 52: 29-37. PDF

32. Pastor, J. M.; Santamaría, S.; Méndez, M.; Galeano, J. (2012). Effects of topology on robustness in ecological bipartite networks. Networks and Heterogeneous Media 7: 429-440. PDF

31. Jara-Guerrero, A.; De la Cruz, M.; Méndez, M. (2011). Seed dispersal spectrum of woody species in South Ecuadorian dry forests: environmental correlates and the effect of considering species abundance. Biotropica 43: 722-730. PDF

30. Teixido, A. L.; Méndez, M.; Valladares, F. (2011). Flower size and longevity influence florivory in the large-flowered shrub Cistus ladanifer. Acta Oecologica 37: 418-421. PDF

29. Álvarez-Lao, D. J.; Méndez, M. (2011). Ontogenetic changes and sexual dimorphism in the mandible of adult woolly mammoths (Mammuthus primigenius). Geobios 44: 335-343. PDF

28. Torices, R.; Méndez, M.; Gómez, J. M. (2011). Where do monomorphic sexual systems fit in the evolution of dioecy? Insights from the largest family of Angiosperms. New Phytologist 190: 234-248. PDF

27. Prieto-Benítez, S.; Méndez, M. (2011). Effects of land management on the abundance and richness of spiders (Araneae): a meta-analysis. Biological Conservation 144: 683-691. PDF

26. Torices, R.; Méndez, M. (2011). Influence of inflorescence size on sexual expression and female reproductive success in a monoecious species. Plant Biology 13 (suppl. 1): 78-85. PDF

25. Torices, R.; Méndez, M. (2010). Fruit size decline from the margin to the center of capitula is the result of resource competition and architectural constraints. Oecologia 164: 949-958. PDF

24. García-Verdugo, C.; Méndez, M.; Velázquez-Rosas, N.; Balaguer, L. (2010). Contrasting patterns of morphological and physiological differentiation across insular environments: phenotypic variation and heritability of light-related traits in Olea europaea. Oecologia 164: 647-655. PDF

23. Aragón, C. F.; Méndez, M.; Escudero, A. (2010). Linkages among life-history components through time: Carryover effects on the demography of a short-lived polycarpic plant. American Journal of Botany 97: 1492-1497. PDF

22. Méndez, M.; Munzinger, J. (2010). Planchonella, first record of gynomonoecy for the family Sapotaceae. Plant Systematics and Evolution 287: 65-73. PDF

21. García-Camacho, R.; Méndez, M.; Escudero, A. (2009). Pollination context effects in the high-mountain dimorphic Armeria caespitosa (Plumbaginaceae): Neighborhood is something more than density. American Journal of Botany 96: 1620-1626. PDF

20. Aragón, C. F.; Méndez, M.; Escudero, A. (2009). Survival costs of reproduction in a short-lived perennial plant: live hard, die young. American Journal of Botany 96: 904-911. PDF

19. DeSoto, L.; Quintanilla, L. G.; Méndez, M. (2008). Environmental sex determination in ferns: effects of nutrient availability and individual density in Woodwardia radicans. Journal of Ecology 96: 1319-1327. PDF

18. Méndez, M.; García, D.; Maestre, F. T.; Escudero, A. (2008). More ecology is needed to restore Mediterranean ecosystems: a reply to Valladares & Gianoli. Restoration Ecology 16: 210-216. PDF

17. Lázaro, A.; Méndez, M. (2007). Variation in sexual expression in the monoecious shrub Buxus balearica at different scales. Plant Biology 9: 736-744. PDF

16. Méndez, M.; Karlsson, P. S. (2007). Equivalence of three allocation currencies as estimates of reproductive allocation and somatic cost of reproduction in Pinguicula vulgaris. Plant Biology 9: 462-468. PDF

15. Quintanilla, L. G.; de Soto, L.; Jiménez, A.; Méndez, M. (2007). Do antheridiogens act via gametophyte size? A study of Woodwardia radicans (Blechnaceae). American Journal of Botany 94: 986-990. PDF

14. Méndez, M.; Gómez, J. M. (2006). Phenotypic gender in Hormathophylla spinosa (Brassicaceae), a perfect hermaphrodite with tetradynamous flowers, is variable. Plant Systematics and Evolution 262: 225-237. PDF

13. Lázaro, A.; Traveset, A.; Méndez, M. (2006). Masting in Buxus balearica: assessing fruiting patterns and processes at a large spatial scale. Oikos 115: 229-240. PDF

12. Méndez, M.; Karlsson, P. S. (2005). Nutrient stoichiometry in Pinguicula vulgaris: nutrient availability, plant size, and reproductive status. Ecology 86: 982-991. PDF

11. Méndez, M.; Karlsson, P. S. (2004). Between-population variation in size-dependent reproduction and reproductive allocation in Pinguicula vulgaris (Lentibulariaceae) and its environmental correlates. Oikos 104: 59-70. PDF

10. Gutiérrez, D.; Menéndez, R.; Méndez, M. (2004). Habitat-based conservation priorities for carabid beetles within the Picos de Europa National Park, northern Spain. Biological Conservation 115: 379-393. PDF

9. Méndez, M.; Traveset, A. (2003). Sexual allocation in single-flowered individuals of a hermaphroditic species in relation to plant and flower size. Oecologia 137: 69-75. PDF

8. Méndez, M. (2001). Sexual mass allocation in species with inflorescences as pollination units: a comparison between Arum italicum and Arisaema (Araceae). American Journal of Botany 88: 1781-1785. PDF

7. Méndez, M.; Díaz, A. (2001). Flowering dynamics in Arum italicum (Araceae): relative role of inflorescence traits, flowering synchrony, and pollination context on fruit initiation. American Journal of Botany 88: 1774-1780. PDF

6. Méndez, M.; Gwynn Jones, D.; Manetas, Y. (1999). Enhanced UV-B radiation under field conditions increases anthocyanin and reduces the risk of photoinhibition bout does not affect growth in the carnivorous plant Pinguicula vulgaris. New Phytologist 144: 275-282. PDF

5. Méndez, M.; Karlsson, P. S. (1999). Costs and benefits of carnivory in plants: insights from the photosynthetic performance of four carnivorous plants in a subarctic environment. Oikos 86: 105-112. PDF

4. Méndez, M. (1999). Effects of sexual reproduction on growth and vegetative propagation in the perennial geophyte Arum italicum (Araceae). Plant Biology 1: 115-120. PDF

3. Méndez, M. (1998). Modification of phenotypic and functional gender in Arum italicum (Araceae). American Journal of Botany 85: 225-234. PDF

2. Méndez, M. (1997). Sources of variation in seed mass in Arum italicum (Araceae). International Journal of Plant Sciences 158: 298-305. PDF

1. Méndez, M.; Obeso, J. R. (1993). Size-dependent reproductive and vegetative allocation in Arum italicum (Araceae). Canadian Journal of Botany 71: 309-314. PDF

Darnowski, D.; Bauer, U.; Méndez, M.; Horner, J.; Plachno, B. (2018). Prey selection and specialization by carnivorous plants. In: Ellison, A. M.; Adamec, L. (eds.) Carnivorous plants: physiology, ecology, and evolution: 285-293. Oxford University Press, Oxford.

Karlsson, P. S.; Méndez, M. (2005). The resource economy of plant reproduction. En: Reekie, E. G.; Bazzaz, F. A. (eds.) Reproductive allocation in plants: 1-49. Academic Press, Burlington, MA.