Isabel Martínez

Departamental II – DII. 216B
Area of Biodiversity and Conservation
Universidad Rey Juan Carlos
c/ Tulipán, s/n.
E-28933 Móstoles (Madrid)
Spain
Phone: +34 91 488 7183
Fax: +34 91 664 7490
E-mail: isabel.martinez at urjc.es
1991 – BsC Biological Sciences, especiality in Botany (Universidad Complutense de Madrid)
1997 – Ph D in Biological Sciences (Universidad Complutense de Madrid)
Full professor.
I teach Iberian Ecosystems in the degree in Biology, Management of Protected Areas in the degree in Environmental Sciences and Biodiversity and Conservation of Lichens and Bryophytes at the master level.
My main research interests focus on conservation biology of lichens, community ecology of epiphytic and terrestrial lichens, population dynamics of lichens in fragmented landscapes and the effect of climate change on lichens.
Fernández-Prado, N.; Prieto, M.; Martínez, I.; Aragón, G. (In press). Líquenes epífitos de los tepuyes de la cuenca alta del río Nangaritza, cordillera del Cóndor (Ecuador). Caldasia.
Ellis, C. J.; Asplund, J.; Benesperi, R.; Branquinho, C.; DiNuzzo, L.; Hurtado, P.; Martínez, I.; Matos, P.; Nascimbene, J.; Pinho, P.; Prieto, M.; et al. (2021). Functional traits in lichen ecology: A review of challenge and opportunity. Microorganisms 9: 766.
Hurtado, P.; Matos, P.; Aragón, G.; Branquinho, C.; Prieto, M.; Martínez, I. (2020). How much matching there is in functional, phylogenetic and taxonomic optima of epiphytic macrolichen communities along an European climatic gradient? Science of the Total Environment 712: 136533.
Hurtado, P.; Prieto, M.; Martínez-Vilalta, J.; Giordani, P.; Aragón, G.; López-Angulo, J.; Kosuthová, A.; Merinero, S.; Díaz-Peña, E. M.; Rosas, T.; Benesperi, R.; Bianchi, E.; Grube, M.; Mayrhofer, H.; Nascimbene, J.; Wedin, M.; Westberg, M.; Martínez, I. (2020). Disentangling functional trait variation and covariation in epiphytic lichens along a continent-wide latitudinal gradient. Proceedings of the Royal Society B 287: 20192862.
Hurtado, P.; Prieto, M.; Aragón, G.; de Bello, F.; Martínez, I. (2020). Intraspecific variability drives functional changes in lichen epiphytic communities across Europe. Ecology 101: e03017. – Q1, Junio
Aragón, G.; Martínez, I.; Hurtado, P.; Benítez, A.; Rodríguez, C.; Prieto, M. (2019). Using growth forms to predict epiphytic lichen abundance in a wide variety of forest types. Diversity 11: 51.
Cardós, J. L. H.; Prieto, M.; Jylhä, M.; Aragón, G.; Molina, M. C.; Martínez, I.; Rikkinen, J. (2019). A case study on the re-establishment of the cyanolichen symbiosis: where do the compatible photobionts come from? Annals of Botany 124: 379–388.
Concostrina-Zubiri, L.; Arenas, J. M.; Martínez, I.; Escudero, A. (2019). Unassisted establishment of biological soil crusts on dryland road slopes. Web Ecology 19: 39-51.
Hurtado, P.; Prieto, M.; Aragón, G.; Escudero, A.; Martínez, I. (2019). Critical predictors of functional phylogenetic and taxonomic diversity are geographically structured in lichen epiphytic communities. Journal of Ecology 107: 2303-2316.
Rodríguez-Peñate, A. E.; Escudero, A.; Martínez, I.; Madrigal-González, J. (2019). Unveiling annual growth chronologies from inter-nodal branch elongations in a fruticose lichen in southern Europe. Fungal Biology 123: 824-829.
Cardós, J. L. H.; Martínez, I.; Aragón, G.; Ellis, C. J. (2018). Role of past and present landscape structure in determining epiphyte richness in fragmented Mediterranean forests. Landscape Ecology 33: 1757-1768.
Concostrina-Zubiri, L.; Martínez, I.; Escudero, A. (2018). Lichen-biocrust diversity in a fragmented dryland: fine scale factors are better predictors than landscape structure. Science of the Total Environment 628-629: 882-892.
Cardós, J. L. H.; Aragón, G.; Martínez, I. (2017). A species on a tightrope: Establishment limitations of an endangered lichen in a fragmented Mediterranean landscape. American Journal of Botany 104: 527-537.
Merinero, S.; Aragón, G.; Martínez, I. (2017). Intraspecific life history variation in contrasting habitats: Insights from an obligate symbiotic organism. American Journal of Botany 104: 1099-1107.
Merinero, S.; Méndez, M.; Aragón, G.; Martínez, I. (2017). Variation in the reproductive strategy of a lichenized fungus along a climatic gradient. Annals of Botany 120: 63-70.
Otálora, M. A. G.; Martínez, I.; Aragón, G.; Wedin, M. (2017). Species delimitation and phylogeography of the Pectenia species-complex: A misunderstood case of species-pairs in lichenized fungi, where reproduction mode does not delimit lineages. Fungal Biology 121: 222-233. PDF
Prieto, M.; Martínez, I.; Aragón, G.; Verdú, M. (2017). Phylogenetic and functional structure of lichen communities under contrasting environmental conditions. Journal of Vegetation Science 28: 871–881.
Rubio-Salcedo, M.; Psomas, A.; Prieto, M.; Zimmermann, N. E.; Martínez, I. (2017). Case study of the implications of climate change for lichen diversity and distributions. Biodiversity and Conservation 26: 1121-1141. PDF
Aragón, G.; Belinchón, R.; Martínez, I.; Prieto, M. (2016). A survey method for assessing the richness of epiphytic lichens using growth forms. Ecological Indicators 62: 101–105.
Cardós, J. L. H.; Martínez, I.; Calvo, V.; Aragón, G. (2016). Epiphyte communities in Mediterranean fragmented forests: importance of the fragment size and the surrounding matrix. Landscape Ecology 31: 1975-1995. PDF
Aragón, G.; Abuja, L.; Belinchón, R.; Martínez, I. (2015). Edge type determines the intensity of forest edge effect on epiphytic communities. European Journal of Forest Research 134: 443–451. PDF
Matos, P.; Pinho, P.; Aragón, G.; Martínez, I.; Nunes, A.; Soares, A. M. V. M.; Branquinho, C. (2015). Lichen traits responding to aridity. Journal of Ecology 103: 451–458. PDF
Merinero, S.; Martínez, I.; Rubio-Salcedo, M.; Gauslaa, Y. (2015). Epiphytic lichen growth in Mediterranean forests: Effects of proximity to the ground and reproductive stage. Basic and Applied Ecology 16: 220-230. PDF
Otálora, M. A. G.; Belinchón, R.; Prieto, M.; Aragón, G.; Izquierdo, P.; MartínezZ, I. (2015). The threatened epiphytic lichen Lobaria pulmonaria in the Iberian Peninsula: Genetic diversity and structure across a latitudinal gradient. Fungal Biology 119: 802-811. PDF
Prieto, M.; Romera, L.; Merinero, S.; Aragón, G.; Martínez, I. (2015). Development and characterization of fungal specific microsatellite markers in the lichen Lobarina scrobiculata (Lobariaceae, Ascomycota). The Lichenologist 47: 183–186. PDF
Rubio-Salcedo, M.; Merinero, S.; Martínez, I. (2015). Tree species and microhabitat influence the population structure of the epiphytic lichen Lobaria pulmonaria. Fungal Ecology 18: 1-9. PDF
Concostrina-Zubiri, L.; Huber-Sannwald, E.; Martínez, I.; Flores Flores, J. L.; Reyes-Agüero, J. A.; Escudero, A.; Belnap, J. (2014). Biological soil crusts across disturbance–recovery scenarios: effect of grazing regime on community dynamics. Ecological Applications 24: 1863-1877. PDF
Concostrina-Zubiri, L.; Martínez, I.; Rabasa, S. G.; Escudero, A. (2014). The influence of environmental factors on biological soil crust: from a community perspective to a species level. Journal of Vegetation Science 25: 503-513. PDF
Concostrina-Zubiri, L.; Pescador, D. S.; Martínez, I.; Escudero, A. (2014). Climate and small scale factors determine functional diversity shifts of biological soil crusts in Iberian drylands. Biodiversity and Conservation 23: 1757-1770. PDF
Martínez, I.; Flores, T.; Aragón, G.; Otálora, M. A. G.; Rubio-Salcedo, M. (2014). What factors influence the occurrence of the genus Degelia (a threatened lichen) in central Spain? Fungal Ecology 11: 50-59. PDF
Merinero, S.; Rubio-Salcedo, M.; Aragón, G.; Martínez, I. (2014). Environmental factors that drive the distribution and abundance of a threatened cyanolichen in Southern Europe: A multi-scale approach. American Journal of Botany 101: 1876-1885. PDF
Aragón, G.; Belinchón, R.; Martínez, I.; Prieto, M. (2013). Estimating epiphytic lichen richness by simple families in Mediterranean forests. Forest Ecology and Management 310: 187-193. PDF
Concostrina-Zubiri, L.; Huber-Sannwald, E.; Martínez, I.; Flores Flores, J. L.; Escudero, A. (2013). Biological soil crusts greatly contribute to small-scale soil heterogeneity along a grazing gradient. Soil Biology and Biochemistry 64: 28-36. PDF
Otálora, M. A. G.; Aragón, G.; Martínez, I.; Wedin, M. (2013). Cardinal characters on a slippery slope – A re-evaluation of phylogeny, character evolution, and evolutionary rates in the jelly lichens (Collemataceae s. str). Molecular Phylogenetics and Evolution 68: 185-198. PDF
Otálora, M. A. G.; Salvador, C.; Martínez, I.; Aragón, G. (2013). Does the reproductive strategy affect the transmission and genetic diversity of bionts in cyanolichens? A case study using two closely related species. Microbial Ecology 65: 517-530. PDF
Rubio-Salcedo, M.; Martínez, I.; Carreño, F.; Escudero, A. (2013). Poor effectiveness of the Natura 2000 network protecting Mediterranean lichen species. Journal for Nature Conservation 21: 1-9. PDF
Aragón, G.; Martínez, I.; García, A. (2012). Loss of epiphytic diversity along a latitudinal gradient in Southern Europe. Science of Total Environment 426: 188-195. PDF
Belinchón, R.; Martínez, I.; Aragón, G.; Escudero, A. (2012). Lichen species co-occurrence patterns along an edge-interior Mediterranean forest gradient. Acta Oecologica 43: 150-157. PDF
Escolar, C.; Martínez, I.; Matthew, A. B.; Maestre, F. T. (2012). Warming reduces the growth and diversity of biological soil crust in a semi-arid environment: implications for ecosystem structure and function. Philosophical Transactions of the Royal society B 367: 3087-3099. PDF
Martínez, I.; Flores, T.; Otálora, M. A. G.; Belinchón, R.; Prieto, M.; Aragón, G.; Escudero, A. (2012). Multiple scale environmental modulation of lichen reproduction. Fungal Biology 116: 1192-1201. PDF
Prieto, M.; Martínez, I.; Aragón, G.; Gueidan, C.; Lutzoni, F. (2012). Molecular phylogeny of Heteroplacidium, Placidium and related catapyrenoid genera (Verrucariaceae, lichen-forming Ascomycota). American Journal of Botany 99: 23-35. PDF
Belinchón, R.; Martínez, I.; Aragón, G.; Escudero, A.; de la Cruz, M. (2011). Fine spatial pattern of an epiphytic lichen species is affected by habitat conditions in two forest types in the Iberian Mediterranean Region. Fungal Biology 115: 1270-1278.
Castillo-Monroy, A. P.; Bowker, M. A.; Maestre, F. T.; Rodríguez-Echeverría, S.; Martínez, I.; Barraza-Zepeda C. E.; Escolar C. (2011). Relationship between biological soil crust, bacterial diversity and abundance and ecosystem functioning: Insights from a semi-arid Mediterranean environment. Journal of Vegetation Science 22: 165-174. PDF
Maestre, F. T.; Bowker, M. A.; Cantón, Y.; Castillo-Monroy, A. P.; Cortina, J.; Escolar, C.; Escudero, A.; Lázaro, R.; Martínez, I. (2011). Ecology and functional roles of biological soil crusts in semi-arid ecosystems of Spain. Journal of Arid Environments 75: 1282-1291. PDF
Otálora, M. G.; Martínez, I.; Belinchón, R.; Widmer, I.; Aragón, G.; Escudero, A.; Scheidegger, C. (2011). Remnants fragments preserve genetic diversity of the old forest lichen Lobaria pulmonaria in a fragmented Mediterranean mountain forest. Biodiversity and Conservation 20: 1239-1254. PDF
Aragón, G.; López, R.; Martínez, I. (2010). Effects of Mediterranean dehesa management on epiphytic lichens. Science of the Total Environment 409: 116-122. PDF
Aragón, G.; Martínez, I.; Izquierdo, P.; Belinchón, R.; Escudero, A. (2010). Effects of forest management on epiphytic lichen diversity in Mediterranean forests. Applied Vegetation Science 13: 183-194. PDF
Maestre, F. T.; Bowker, M. A.; Escolar, C.; Puche, M. D.; Soliveres, S.; Mouro, S.; García-Palacios, P.; Castillo-Monroy, A. P.; Martínez, I.; Escudero, A. (2010). Do biotic interactions modulate ecosystem functioning along abiotic stress gradients? Insights from semi-arid Mediterranean plant and biological soil crust communities. Philosophical Transactions of the Royal Society B 365: 2057-2070. PDF
Otálora, M. A. G.; Martínez, I.; O´Brien, H.; Molina, M. C.; Aragón, G.; Lutzoni, F. (2010). Multiple origins of high reciprocal symbiotic specificity at an intercontinental spatial scale among gelatinous lichens (Collemataceae, Lecanoromycetes). Molecular Phylogenetics and Evolution 56: 1089-1095. PDF
Otálora M. A. G.; Martínez, I.; Aragón, G.; Molina, M. C.; Lutzoni, F. (2010). Disentangling the Collema-Leptogium complex through a molecular phylogenetic study of the Collemataceae (Peltigerales, lichen-forming Ascomycota). Mycologia 102: 279-290. PDF
Otálora, M. A. G.; Martínez, I.; Aragón, G.; Molina, M. C. (2010). Phylogeography and divergence date estimates of a lichen species complex with a disjunct distribution pattern. American Journal of Botany 97: 216-223. PDF
Prieto, M.; Martínez, I.; Aragón, G.; Otálora, M. A. G.; Lutzoni, F. (2010). Phylogenetic study of Catapyrenium s. str. (Verrucariaceae, lichen-forming Ascomycota) and related genus Placidiopsis. Mycologia 102: 291-304. PDF
Prieto, M.; Aragón, G.; Martínez, I. (2010). The genus Catapyrenium s. lat. (Verrucariaceae) in the Iberian Peninsula and the Balearic Islands. The Lichenologist 42: 637-684. PDF
Prieto, M.; Martínez, I.; Aragón, G. (2010). The genus Placidiopsis in the Iberian Peninsula and the Balearic Islands. Mycotaxon 114: 463-472. PDF
2009
Belinchón, R.; Martínez, I.; Otálora, M. A. G.; Aragón, G.; Dimas, J.; Escudero, A. (2009). Fragment quality and matrix affect epiphytic performance in Mediterranean forest landscape. American Journal of Botany 96: 1974-1982. PDF
Maestre, F. T.; Bowker, M. A.; Puche, M. D.; Hinojosa, M. B.; Martínez, I.; García-Palacios, P.; Castillo, A. P.; Soliveres, S.; Luzuriaga, A. L.; Sánchez, A. M.; Carreira, J. A.; Gallardo, A.; Escudero, A. (2009). Shrub encroachement can reverse desertification in semi-arid Mediterranean grasslands. Ecology Letters 12: 930-941. PDF
Maestre, F. T.; Martínez, I.; Escolar, C.; Escudero, A. (2009). On the relationship between abiotic stress and co-occurrence patterns: an assessment at the community level using soil lichen communities and multiple stress gradients. Oikos 118: 1015-1022. PDF
2008
Maestre, F. T.; Escolar, C.; Martínez, I.; Escudero, A. (2008). Are soil lichen communities structured by biotic interactions? A null model analysis. Journal of Vegetation Science 19: 261-266. PDF
Otálora, M. A. G.; Martínez, I.; Molina, M. C.; Aragón, G.; Lutzani, F. (2008). Phylogenetic relationships and taxonomy of the Leptogiumlichenoides group (Collemataceae, Ascomycota) in Europe. Taxon 57: 907-921. PDF
Prieto, M.; Aragón, G.; Martínez, I. (2008). New records in the lichen family Verrucariaceae (Ascomycota) from Argentina. Boletín de la Sociedad Argentina de Botánica 43: 205-210. PDF
Prieto, M.; Aragón, G.; Martínez, I.; Breuss, O. (2008). A new species of Anthracocarpon (Verrucariaceae) from Argentina. The Briologist 111: 128-132. PDF
2007
Belinchón, R.; Martínez, I.; Escudero, A.; Aragón, G.; Valladares, F. (2007). Edge effects on epiphytic communities in a Mediterranean Quercus pyrenaica forest. Journal of Vegetation Science 18: 81-90. PDF
Escudero, A.; Martínez, I.; de la Cruz, A.; Otálora, M. A. G.; Maestre, F. T. (2007). Soil lichens have species-specific effects on the seedling emergence of three gypsophile plant species. Journal of Arid Environments 70: 18-28. PDF
2006
Martínez, I.; Carreño, F.; Escudero, A.; Rubio, A. (2006). Are threatened lichen species well-protected in Spain? Effectiveness of a protected areas network. Biological Conservation 133: 500-511. PDF
Martínez, I.; Escudero, A.; Maestre, F.; de la Cruz, A. ; Guerrero, C.; Rubio, A. (2006). Small scale abundance of mosses and lichens forming soil biological crusts in two semi-arid gypsum environments. Australian Journal of Botany 54: 339-348. PDF
2005
Aragón, G.; Otálora, M. A. G.; Martínez, I. (2005). New data of the genus Leptogium (lichenized Ascomycetes) in the Iberian Peninsula. Nova Hedwigia 80:199-226.
Maestre, F. T.; Escudero, A.; Martínez, I.; Guerrero, C.; Rubio, A. (2005). Does spatial pattern matter to ecosystem functioning? Insights from biological soil crusts. Functional Ecology 19: 566-573. PDF
2004
Aragón, G.; Martínez, I. , Otálora, M. A. G. (2004). Leptogium subaridum, a new Mediterranean-SW American disjunction. The Lichenologist 36: 163-165.
Aragón, G.; Martínez, I. , Otálora, M. A. G. (2004). New data on the distribution of Leptogium azureum (Swartz) Mont. The Lichenologist 36: 345-347.
Aragón, G.; Sarrión, F. J.; Martínez, I. (2004). Epiphytic lichens on Juniperus oxycedrus in the Iberian Peninsula. Nova Hedwigia 78: 45-56.
Burgaz, A. R.; Ahti, T.; Martínez, I. (2004). Contribution to the study of the genus Cladonia in the Iberian Peninsula. I. Supergroup Crustaceae. Nova Hedwigia (en prensa).
Muñoz, J.; Felicísimo, A. M.; Cabezas, F.; Burgaz, A. R.; Martínez, I. (2004). Wind as a long-distance dispersal vehicle in the Southern Hemisphere. Science 304: 1144-1147. PDF
Martínez, I. , Aragón, G. (2004). The Lecanora varia group in Spain: species with amphitecial cortex. The Bryologist 107: 222-230. PDF
Otálora, M. A. G.; Aragón, G.; Martínez, I.; Molina, M.C. (2004). A new corticolous species of Leptogium (Collemataceae) from Spain. The Lichenologist 36
2003
Aragón, G.; Martínez, I. (2003). Bacidia iberica, a new lichen from Spain, and its relationship to Bacidia rubella. The Bryologist 106: 143-146.
Martínez, I.; Aragón, G. (2003). Ophioparma juniperiicola, a new lichen from Spain. The Bryologist 106: 528-531.
Martínez, I.; Aragón, G.; Sarrión, F. J.; Escudero, A. , Burgaz, A. R. ,Coppins, B. (2003). Threatened lichens in central Spain. Cryptogamie, Mycologie 24: 73-97.
Martínez, I.; Burgaz, A. R.; Vitikainen, O.; Escudero, A. (2003). Distribution pattern of the genus Peltigera Willd. The Lichenologist 35: 301-323. PDF
2002
Aragón, G.; Martínez, I. (2002). Candelariella faginea and C. viae-lacteae, new to SW Spain. The Lichenologist 34: 81-88.
Burgaz, A. R.; Martínez, I. (2002). The Placynthium nigrum group in the Iberian Peninsula. Cryptogamie, Mycologie 23: 367-375.
Martínez, I. (2002). Lichenicolous fungi from the Iberian Peninsula and the Macaronesian area. Nova Hedwigia 74: 51-67.
Martínez, I.; Aragón, G.; Burgaz, A R. (2002). Epiphytic lichens and lichenicoloud fungi from “Serranía de Cuenca” mountains (“Sistema Ibérico”, Cuenca Province, Spain). Herzogia 15: 37-49.
1993-2001
Burgaz, A. R.; Martínez, I. (2001). The genera Koerberia, Leptochidium, Massalongia and Polychidium (Peltigerales s. l.) in the Iberian Peninsula. Nova Hedwigia 73: 381-392.
Aragón, G. , Martínez, I. (1999). Contribución al conocimiento de los líquenes epifíticos de la Sierra de Alcaraz (Albacete, España). Cryptogamie, Mycologie 20: 57-70.
Aragón, G.; Martínez, I.; Burgaz, A. R. (1999). Contribución al conocimiento de los líquenes epifíticos del valle de Sanabria (Zamora). Cryptogamie, Mycologie 20: 137-152.
Burgaz, A. R.; Martínez, I. (1999). The genus Nephroma Ach. in the Iberian Peninsula. Cryptogamie, Mycologie 20: 225- 235.
Martínez, I.; Aragón, G.; Lumbsch, H. T. (1999). Lecanora paramerae, a new lichen from Spain. The Lichenologist 31: 315-318.
Martínez, I.; Hafellner, J. (1998). Lichens and lichenicolous fungi growing on Peltigerales in the Iberian Peninsula and the Canary Islands. Mycotaxon 69: 271-310.
Aragón, G.; Martínez, I. (1997). Contribución al conocimiento de los líquenes epifíticos de los Montes de Toledo (Toledo, España). Cryptogamie, Bryol. Lichénol. 18: 63-75.
Martínez, I.; Burgaz, A. R. (1997). Additions to the chemistry, ecology and distribution of Peltigera melanorrhiza and P. neopolydactyla. Mycotaxon 63: 389-394.
Martínez, I.; Burgaz, A. R. (1997). Revision of the genus Solorina (Lichenes) in Europe based on spore size variation. Annales Botanici Fennici 35: 137-142.
Martínez, I.; Burgaz, A. R.; Vitikainen, O. (1997). Studies on the genus Peltigera in the Iberian Peninsula. II. Nova Hedwigia 64: 111-135.
Martínez, I.; Aragón, G. (1996). Líquenes epifíticos de la vertiente norte del Puerto de la Quesera, Macizo de Ayllón (Segovia, España). Cryptogamie, Bryol. Lichénol. 1996: 143-156.
Martínez, I.; Burgaz, A. R. (1996). Anatomical study of Peltigera canina, P. membranacea and P. praetextata (Lichenized, Ascomycotina). Annales Botanici Fennici 33: 223-229.
Martínez, I.; Burgaz, A. R. (1994). Estudio de la anatomía del talo de Peltigera canina y P. membranacea. Cryptogamie, Bryol. Lichénol. 15: 321-336.
Martínez, I.; Burgaz, A. R. (1993). Estudio del género Peltigera en España peninsular. I. Cryptogamie, Bryol. Lichénol. 14: 341-352.
Sarrión, F. J.; Martínez, I.; Burgaz, A. R. (1993). Líquenes epífitos de Sierra Madrona (Ciudad Real, España). Cryptogamie, Bryol. Lichénol. 14: 389-400.
Otros trabajos relevantes
Martínez, I. (2016) Los líquenes y su conservación en España. Conservación Vegetal 20. PDF
Concostrina-Zubiri, L.; Martínez, I.; Huber-Sannwald, E.; Escudero, A. (2013). Efectos y respuestas de la costra biológica del suelo en ecosistemas áridos: avances recientes a nivel de especie. Ecosistemas 22: 95-100. PDF
Martínez, I.; Belinchón, R.; Otálora, M. A. G.; Aragón, G.; Prieto, M.; Escudero, A. (2011). Efectos de la fragmentación de los bosques sobre los líquenes epífitos en la Región Mediterránea. Ecosistemas 20: 54-67. PDF
Burgaz, A. R.; Martínez, I.; Aragón, G. (2007). Criptógamas: briófitos y líquenes de Teruel. Teruel 90: 165-208.
Burgaz, A. R.; Martínez, I. (2003). Peltigerales. Lobariaceae, Nephromataceae, Peltigeraceae. Flora Liquenológica Ibérica, vol. 1. Universidad de Murcia, Murcia.
Aragón, G.; Martínez, I.; Burgaz, A. R. (2001). Macrolíquenes de Castilla-La Mancha. Cuadernos de Estudios Manchegos: 1-190. Junta de Castilla-La Mancha, Ciudad Real.
Martínez, I. (1999). Taxonomía del género Peltigera Willd. (Ascomycetes liquenizados) en la Península Ibérica y estudio de sus hongos liquenícolas. Monografías de RUIZIA 15: 1-200. CSIC, Madrid.